is calcification a co2 source
yes
selective sweeps affect:
diversity
LD (linkage disequilibriu,) —> also breaks down with physical distance
differentiation
LD
Linkage disequilibrium
non-random association of alleles at two or more loci
Two homologous chromosomes in diploids (except
in the gametes, which are haploid)
Each chromosome has two (identical) sister
chromatids
Recombination:
• Chromosomes exchange short segments
• happens in the germline during meiosis.
Proteomics
is the large-scale study of proteins. The proteome is the entire set of proteins that is produced or modified by an organsism or system at the time-point of sampling.
Metabolomics
is the study of small molecules, commonly known as metabolites, within cells, biofluids, tissues or organisms. Collectively, these small molecules and their interactions within a biological system are know as the metabolome.
Genomics
– the systematic study of an organsism’s genome.
Transcriptomics
– the transcriptome is the total complement of ribonucleic acid (RNA) transcripts in a cell and consists of coding messenger and non-coding ribosomal, transfer, small nuclear, small interfering, micro and long-non-coding RNAs. Trancriptomics – is the analysis of mRNAs and provides direct insight into cell- and tissue-specific genen expression features.
Metaproteomics
Metaproteomics [complement of proteins synthesized by an organism/community of (micro)organisms present in an environmental sample] - not all proteins are detected
coverage per organism depends on the representation of that organism in the community
abundance of the protein in the cell
methodological factors (e.g. polarity)
the metaproteome informs about the proteins synthesized by a microbial community at the time of sampling
(A) Increase MPA resilience
(ability of the
ecosystem to resist, recover and adapt to
climate change while maintaining ecosystem
functions and services; Holling 1973)
-
Protect critical habitat areas (ecologically
important e.g. fish spawning areas and
nesting sites and climate refugia)
Maintain connectivity (larval dispersal, mobile
species, links between different habitats)
Maintain ecosystem function
Reduce other stressors (e.g. fishing)
Use ecosystem based management
levels of biol interactions and ecology subdisciplines:
ecophysiology (ind. org.)
pop ecology
community ecology (interacting species)
ecosystem ecology (entire ecosystem: matter and energy fluxes)
biogeochemistry (entire earth at ecosystem)
species either structured between —> 2 major life istory trade offs
age: k stratege (biomass)
kelp
size: (reproduction) r stratege —> growth rate
diatoms
formula pop growth
Nt = No * (1+R)^t
processes determining k (biomass)
plant nutrients
food items
space availability
sexual reproduction distinction:
iteroparity
sveral repr through life
macroalgae, invertebrates, mar vertebrates
semelparity: one production, them death
cephalopods, salmon
relevant for matter fluxes
total resources to organism:
fecundity (nr of sperms produced)
growth
maintainance
defence/longevity
comm assembly as step wise filtering process
global sp pool
translocation filter
introduced species pool
dispersal filter
regional species pool
abiotic filter
habitat sp pool
biotic filter
biotic and abiotic = ecological niche
why can so many species coexist
many dimensions of environmental factors
competition trade of
high max growth rate or
live on low resources (better competitor)
marine vs terrestrial food chains
size interval in terestrial much shorter (in water higher length distrb)
smaller differences and no size trend in terrestrial (big and small herbivores)
water: consumer feed over prey —> increasing size (exc. wales, parasites)
complexity in food chain
ML
jelly food chain
not only predator/prey—> parasitism mixotrophy etc.
up to 5TL in 1TL (pred/prey can range from 1:1 to 15:1)
concept for food web
length
stablity
food web s in food web
comlexity interactions beyond normal idea
measuring TL
amount of energy
longest position
shortest position
low productivity resulting in ___ efficiency
higher
more TL because organisms tend to be smaller bc shows advantage to get more nutrients
more smaller organisms, more TL
more TL —> higher EE
—>
high nutrients = less stepways and therefore lower EE
—> explains the lower div. in higher productivity
quantification of animal diet
gut content
quantitative pcr methods
stable isotope analysis
hypotheses to determine TL limitation
energy contraint hyp: declined
low prod is not food chain shortening
ecosystem size hyp:
larger org = higher TL —> lower metabolic rate and less energy uptake
productive space hyp: same size but more prod= more diverse
dynamic energy budget
food —> storage
via growth t structure
via somatic maintenance to met. rate
via meturity m. to met. work
via repr maturation to gonads
most important for energy cell budget
pbs
ion regulation
UCP
uncoupling proteins: release protons into mt matrix without atp production
avoid accumulation of O radicals
generate heat (brown adipose tissue)
—> activated by proton leak
most important tight junctions
clauding
occluding
feedback loops providing homeostasis
—> activate ion transport to maintain ph
loops to control body temp (homeothermic animals—> loose or keep heat while opening pasage)
loop to maintain ph in cell: enzymatic reaction sensitiv eto ph change (glycolysis)
processes contributing to cellular miantenance:
feedback loop
diffusion
compartimentalization of substances
buffering of atp conc via phosphagen systems
macromolecule repair
immune defense
detoxification of damaged macromolecules
signalling processes (hormonal, neural)
organic osmolyte regulation
how can mar org maintain homeostasis
adaptation (mutation, recomb, plastiticty/acclimatisation)
epigenetics ( modification of chrmatin/dna, also plasticity)
processes affected by thermal acclimatisation
gene expression/pbs
enzyme activity
ultrastructural modification
heat and metabolic rates
points against equilibrium
inbreeding
assortative mating
migration
nat selection
pop structure
fixation index:
how much reduction is expected at heterozygosity (is relative to HW eq)
<Fis: broadcast spawning, plamctonic larvae
>Fis: brooders, internal fertilization, spermcastings
allele freq change over tme bc of
drift
selection
—> evolution
effective pop size influenced by
change in pop size
varriation in repr success
uneven breeding sex ratios
why are Ne/N low in marine pop?
sweeptake repr system
population bias
fluctuation in pop size
assisted evolution on corals
induce acclimatizatiob
modification of microbial symb communities
selective breeding
evolution of symbiondinium
mechanisms regulation diversity
regional
—> immigration, sp sorting/speciation
<— extinction, dispersal
local
—> dispersal
<— emigration, random ext., competition
—> and <— loc sp interactions, consumption pred.
extinction drivers
indirect:
conflicts
epidemics
demographic
economic etc
direct
invasive sp
pollution
climate change
direct exploitation (trawl, hunt)
land/sea use change
metagenomics overview
genomics (genome)
transcriptomics (mRNA)
proteomics (all proteins ecpressed in tissue)
metabolomics (global metabolite system)
Age of DOC
labile:
minutes-days
0.2 Gt
semi-labile
weeks-months
6 Gt
semi-refractory
years-cent
50 GT
refractory:
cent-millenia
660 Gt (anja 630)
DOC sources and sinks
source
direct exudation
viral lysis
pom degradation
sinks:
bacterial consumption/degradation
aggregation
opal at sedoments
atlantic
low to moderate fuxes
high dissosolution rate due to low deep Si(OH)4
pacific
low to high fluxes
low diss due to high deep sioh4
southern ocean
high fluxes
high deep sioh4 allow opal presentation
biomineralisation ocean supply
weathering on land
transport on rivers
deposition in ocean
major source of CO2 in ML
bacteria responsible for most of respiration (OC—>IC)
60% is <1µm
ML link
active biomass: 40-70% living C reservoir
—> production of living BM from dead BM (detritus,DOM)
diverting biomass to higher TL
BGE increase/decrease
increses with incresing phyt prod
decrease with increasing C:N ratios of substrates
varies with comm structure
determined by processes like catabolism, anabolism, atp storage
selfish uptake mechanism
profitable for free living cell
take up large oligomers which are then degraded in periplasmatic space = minimizing hydrolysate loss
main components of biol pump
gravitational pump
sinking speed of different org:
mesopelagic migrant pump —> diel vertical migration
avoidance of optically oriented predators
phys constraints important (hypoxia tolerance)
organism target certain isolumen (depth of light level)
flux attenuation
microbial activity
poliquetszooplankton feeding on aggregates
particle fragmention
eddies with high reproduction rate:
cyclonic eddies: anticlockwise at NH
sea level depression and uplifting of isopicnals
upwelling leads ti more productiveness
less srable than anticyclonic
b
bsp: peoebicus sp. home at AMCE bc of O2 anomally there
competitive exclusion principle
nr of species </= number of resources
many dfferent phyt sp coexist —> eddies might be the answer
external signals acclimating to certain seasons
light (sensor of day length)
changes in metabolic rate
physiological indicators of thermal stress
level of ubiquinated proteins
ub proteasome pathways rids cell of denatured proteins
use ub to conjugate spec antibodies
level of HSP
rescue proteins
steps denaturation (ubiqunation) and protein protection (HSP)
native protein unfolding
Ub:
toxic aggregation
ubiquination
proteolysis
HSP
HSP70 binds hydrophobic regions
protein recovery
3 phases of CRSP/CAS
acquisition
working phase
interference phase
solution for fishing down food web
use omnivorous fish
use lower TL (farming down food chain) —> mariculture —> alleviates coastal eutrophication
conceptual model of invasion:
stage 0: source region
filter: uptake (propagle pressure, physio-chemical tolerance, biol interactions)
stage 1 transportation
transport/release (55% ship ballast, 18% unknown
stage 2: introduction
colonization/reproduction( same as first filter)
stage 3: establishment
transport vectors:
intentional
food and games
garden ornamentation
biocontrol
unntentional
aquaculture
planes
ships
steps for MPA design
increase MPA resilience
protokol climate refugua
protect future habitat
increase connectivity
increase heterogeneity
reduce other stressors: minimize cumulative impacts
other methods: dynamic MPAs
blue carbon
sedimentation of org particles
roots/rhizomes/dead leaves of seagrass = OC persistence for centuries in anoxic sediments
seagrass loss due to
nutrient pollution
warming
techniques coping with seagrass loss
single shooting technique
enemy free space
seed based methods/restauration
co benefits seagras
fisheries
water quality
reduction of harmfull bacteria
biodiversity
coastal protection
enhancing baltic blue carbon through:
upscaling from ha to sqkm
enhance thermal tolerance
enhance public awareness
increase MPA to 30%
enable assisted migration/assisted evolution
Last changeda year ago